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Biblidinae
Biblidinae

Biblidinae is the name for a subfamily of nymphalid butterflies that includes the tropical brushfoots. This subfamily was sometimes merged within the Limenitidinae, but they are now recognized as quite distinct lineages. In older literature, this subfamily is sometimes called Eurytelinae.

As of 2008, some 340 valid species are in this subfamily, placed in 38 genera. Most species of Biblidinae are Neotropical, but there are some Old World species and genera in the tribes Biblidini and Epicaliini.

Systematics


The Biblidinae are a taxonomically stable monophyletic group, at least since the "wastebin genus" Catagramma was dismantled around 1950. Six tribes are recognized, and the monophyly of them all is likewise well supported; radical changes in taxonomy and systematics are not expected in this subfamily. The sister group to the Biblidinae is probably the Apaturinae and/or Nymphalinae.

The tribes, in the presumed phylogenetic sequence and with notable genera also listed here, are:

Biblidini Boisduval, 1833

  • Biblis Fabricius, 1807 (= Zonaga)
  • Ariadne Horsfield, 1829 (= Ergolis)
  • Laringa Moore, 1901
  • Eurytela Boisduval, 1833
  • Neptidopsis Aurivillius, 1898
  • Mesoxantha Aurivillius, 1898
  • Byblia Hübner, 1819 (= Hypanis Boisduval, 1833 (non Pander in Menetries, 1832: preoccupied))
  • Mestra Hübner, 1825 (= Cystineura)
  • Archimestra Munroe, 1949
  • Vila Kirby, 1871 (= Lonia, Neptis Hübner, 1819 (non Fabricius, 1807: preoccupied), Olina Doubleday, 1848 (non Robineau-Desvoidy, 1830: preoccupied))

Epicaliini Guenée, 1865

Ageroniini Doubleday, 1847

  • Hamadryas – cracker butterflies, calico butterflies
  • Ectima Doubleday, 1848
  • Panacea Godman & Salvin, 1883 (= Pandora Doubleday, 1848 (non Lamarck, 1799: preoccupied))
  • Batesia Felder & Felder, 1862

Epiphilini Jenkins, 1987

  • Asterope Hübner, [1819]
  • Pyrrhogyra Hübner, 1819 (= Corybas)
  • Epiphile Doubleday, 1844
  • Lucinia Hübner, 1823 (= Autodea)
  • Bolboneura Godman & Salvin, 1877
  • Temenis Hübner, 1819 (= Callicorina, Paromia Hewitson, 1861 (non Westwood, 1851: preoccupied))
  • Nica Hübner, 1826 (= Pseudonica)
  • Peria Kirby, 1871 (= Pelia Doubleday, 1849 (non Bell, 1836: preoccupied))

Eubagini Burmeister, 1878

  • Dynamine Hübner, 1819 (= Arisba, Eubagis, Sironia)

Callicorini Orfila, 1952 – eighty-eights and relatives

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